Hamilton's rule

E634418

Hamilton's rule is a principle in evolutionary biology that explains how altruistic behavior can evolve when the genetic benefits to related individuals, weighted by relatedness, exceed the costs to the actor.

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Predicate Object
instanceOf evolutionary principle
rule in evolutionary biology
theoretical model of altruism
addresses conditions for the evolution of helping behavior
genetic basis of social behavior
appliesTo genetically related individuals
assumes costs and benefits are measured in genetic fitness
weak selection in many derivations
basedOnConcept inclusive fitness
kin selection
category biological rule
centralConcept coefficient of relatedness
cost–benefit analysis of behavior
coreFormula rb > c
definesVariable b (benefit to recipient)
c (cost to actor)
r (coefficient of relatedness)
explains conditions for the spread of altruistic genes
evolution of altruism
kin-directed altruistic behavior
field behavioral ecology
evolutionary biology
population genetics
sociobiology
formulatedInYear 1964
hasGeneralization inclusive fitness maximization frameworks
multiplayer Hamilton's rule
neighbor-modulated fitness models
hasLimitation applies most directly to additive genetic effects
simplest form assumes pairwise interactions
historicalSignificance foundation of inclusive fitness theory
landmark in the study of social evolution
influenced evolutionary psychology of social behavior
modern sociobiology
theoretical work on social evolution
mathematicallyExpressedAs ΔW > 0 if rb − c > 0
namedAfter W. D. Hamilton NERFINISHED
oftenIllustratedBy alarm calling in social mammals
altruism in eusocial insects
food sharing among kin
proposedBy W. D. Hamilton NERFINISHED
publishedIn Journal of Theoretical Biology NERFINISHED
relatedTo inclusive fitness theory
kin selection theory
statesThat an altruistic trait can spread when rb exceeds c
usedFor analyzing social behavior in animals
modeling evolution of cooperation among relatives
predicting when altruistic behaviors evolve

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William D. Hamilton knownFor Hamilton's rule